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Are the covenants carnivores?

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Brutes are mentioned in the books as consuming the meat of 'thorn beasts'. It is also alluded to that they will eat PoW's (marines) that they capture. Grunts however are apparently fed from something called a food nipple, some sort of nutritional sludge.

I would say it would depend on the species of covenant as they are all alien to each other as well as humans.

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